I can provide some first hand experience with how the fuel gauge in the dash reads and how large each tank is. I recently bought an 04’ 2500 and the neck of the rear fuel tank was cracked. My first fuel up attempt lead to gasoline just dumping on the ground instead of going into the truck. I ended up cutting the hose leading to that tank and plugging it off so I could at least put gas in the truck. So now I just have the main/front tank operational only.The rear tank is pretty easy. You'll have to drop the tire, remove the spare tire bracket, then the two tank strap bolts. When disconnecting the fuel filler hoses, I found it easier to remove them from the big tank side, and then just unbolt the fuel filler neck and ground strap from the filler door, it's only three small bolts- easier to get to. Then of course unhook your fuel pump wires and fuel lines when you can lower it enough. Also, while you are at it. I recommend removing the anti siphon flapper valve from the plastic T fitting on top of the little tank, it's the one both filler hoses attach to, you'll see it wobbling in there. It's easy to over tighten those hose clamps and deform it. Do that and filling that thing up will be a nightmare. Ask how I know, had to drop the tank a bit to get that thing out again. I slid the whole flapper assembly out, removed the metal pin and flapper- it's about the size of a quarter, and put the rest back in the T fitting- I felt the T was kinda flimsy without it.
I'm not sure about the gauge reading. There is a float and sender on that pump, I don't know if it is only to shut the pump off when it's empty, or if the truck uses that reading to calculate remaining fuel. I really have no idea.
When I fill the truck up all the way the fuel gauge in the dash only reads 3/4. To verify the main tank was actually filling 100% I fueled up until gas overflowed out of the filler neck. The truck definitely calculates how much fuel is in each tank and then displays a single reading on the dash. Furthermore, I have HPtuners and licensed the truck. The PCM file shows the main tank is 25 gallons and the rear tank is 13 (38 total). Additionally, 25 is 65% of 38 which closely matched my dash read out. Even further I “tuned out”/deleted my rear fuel tank in the PCM file to see if I could get my fuel gauge to read correctly for only using the main tank. I was successful and now when I gas up I have an accurate fuel reading. I usually fill up at a 1/2 tank and I routinely put in 12-13 gallons.
Summary: fuel gauge takes a reading from both tanks and a single reading is calculated by the PCM. The main tank appears to be 25 gallons and rear appears to be 13 gallons.
I will be attempting to replace my fuel lines and the rear tank in the next few days. I’ve found full replacement fuel line kits for 02-03 trucks but not 04 and newer. Does anyone know if the 02-03 kit would work? I reached out to fine lines / sstubes.com and they didn’t think their 02-03 kit would work on an 04 because the evap and emissions systems allegedly were changed in 04. Any insight would be helpful. Thanks
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